Sweelinck did not write for pedals (though I know of at least one exception to this 'rule'). Karges, however adapted them for the use of pedals. At least that's what I think, given the distance between bass voice and tenor voice in some spots. Yet the manuscript does not generally indicate 'pedals' or 'manual' (though there are, again, several exceptions to this 'rule'). So, it's a editorial decision and you are free to play the bass voice manualiter.